Nanami Momozono is alone and homeless after her dad skips town to evade his gambling debts and the debt collectors kick her out of her apartment. So when a man she's just saved from a dog offers her his home, she jumps at the opportunity. But it turns out that his place is a shrine, and Nanami has unwittingly taken over his job as a local deity!

Himemiko, ayakashi ruler of Tatara swamp, is happily dating her human sweetie Kotaro. But yokai/human relationships are hard to manage, especially when the human has no idea who he's dating! When the supernatural starts to intrude, can Himemiko tell Kotaro the truth about herself? Or will she lose him no matter what she does?