Nanao used to be a house cat, but after tragedy struck, he was abandoned on the rough streets of Tokyo. Now, alongside his best friend and fellow stray Machi, Nanao has to fight every day just to get by. Aside from foraging for food and finding somewhere to sleep, Nanao and Machi must remain vigilant about all sorts of dangers, including humans who want stray cats gone.

When a woman named Yoshino starts feeding the neighborhood cats, Nanao reluctantly bonds with her. But the woman has emotional scars and a past entwined with Nanao's. Can Nanao help her heal, and perhaps heal himself in the process?