For more than seven hundred years, the vast, rambling Trelawney Castle in Cornwall--turrets, follies, a room for every day of the year, four miles of corridors and 500,000 acres--was the magnificent and grand "three dimensional calling card" of the earls of Trelawney. By 2008, it is in a complete state of ruin due to the dulled ambition and the financial ineptitude of twenty-four earls, two world wars, the Wall Street crash, and inheritance taxes. Still: the heir to all of it, Kitto, his wife, Jane, their three children, their dog, Kitto's ancient parents, and his aunt Tuffy Scott, an entomologist who studies fleas, all manage to live there and (barely) keep it going.

Four women dominate the story: Jane; Kitto's sister, Blaze, who left Trelawney and made a killing in finance in London, the wildly beautiful, seductive, and long-ago banished Anastasia and her daughter, Ayesha. When Anastasia sends a letter announcing that her nineteen-year-old daughter, Ayesha, will be coming to stay, the long-estranged Blaze and Jane must band together to take charge of their new visitor--and save the house of Trelawney. But both Blaze and Jane are about to discover that the house itself is really only a very small part of what keeps the family together.