Smith introduces a compelling comparative theory to explain variation in the presence of dynasties across democracies and political parties. Drawing on extensive legislator-level data from twelve democracies and detailed candidate-level data from Japan, he examines the inherited advantage that members of dynasties reap throughout their political careers--from candidate selection, to election, to promotion into cabinet. Smith shows how the nature and extent of this advantage, as well as its consequences for representation, vary significantly with the institutional context of electoral rules and features of party organization. His findings extend far beyond Japan, shedding light on the causes and consequences of dynastic politics for democracies around the world.