Young architect Wren Fontaine lands her dream job: restoring Greenleaf House, New York's finest Gilded-Age mansion, to its glory days. But old homes have old secrets: Stephen Greenleaf-heir to what's left of his family's legacy-refuses to reveal what his plans are once the renovation is completed. And still living in a corner of the home is Stephen's 90-year-old Aunt Agnes who's lost in the past, brooding over a long-forgotten scandal while watching Wren with mistrust.

Wren's job becomes more complex when a shady developer who was trying to acquire Greenleaf House is found murdered. And after breaking into a sealed attic, Wren finds a skeleton stuffed in a trunk. She soon realizes the two deaths, a century apart, are strangely related. Meanwhile, a distraction of a different kind appears in the form of her client's niece, the beautiful and seductive Hadley Vanderwerf. As Wren gingerly approaches a romance, she finds that Hadley has her own secrets.

Then a third murder occurs, and the introverted architect is forced to think about people, and about how ill-fated love affairs and obsessions continue to haunt the Greenleafs. In the end, Wren risks her own life to uncover a pair of murderers, separated by a century but connected by motive. She reveals an odd twist in the family tree that forever changes the lives of the Greenleafs, the people who served them, the mansion they all called home-and even Wren herself.